Systematic and synonymic notes on Ceutorhynchinae, with lectotype and neotype designations, and descriptions of three new genera (Coleoptera, Curculionidae)

Publication Type:Journal Article
:1998
Authors:E. Colonnelli
Journal:Fragmenta Entomologica
Volume:30
Pagination:105-175
Type of Article:Article
:0429-288X
:Acallodes saltoides, Amicroplontus arcuatus, Aoxyonyx, Auleutes epilobii ., Barioxyonyx, Brachiodontus, Brevicoeliodes, Ceuthorhynchideus minimus, Ceuthorhynchidius porcellus, Ceuthorrhynchus cingulatus, Ceuthorrhynchus gibbicollis, Ceuthorrhynchus pudicus, Ceuthorrhynchus sainteclairei, Ceutorhynchinae, Ceutorhynchus, Ceutorhynchus abchasicus var. tyli, Ceutorhynchus albicans, Ceutorhynchus assimilis var. major, Ceutorhynchus carnifrons, Ceutorhynchus chalibaeus, Ceutorhynchus cochleariae, Ceutorhynchus contractus, Ceutorhynchus distans, Ceutorhynchus erythropus, Ceutorhynchus fallax, Ceutorhynchus gallorhenanus, Ceutorhynchus granipennis, Ceutorhynchus granulicollis, Ceutorhynchus grenieri, Ceutorhynchus griseus, Ceutorhynchus ilvensis, Ceutorhynchus insecabilis, Ceutorhynchus kaufmanni, Ceutorhynchus minutus, Ceutorhynchus moguntiacus, Ceutorhynchus nanus, Ceutorhynchus nigrostriatus, Ceutorhynchus obesulus, Ceutorhynchus pandellei, Ceutorhynchus picitarsis, Ceutorhynchus pictus, Ceutorhynchus primevus, Ceutorhynchus quadripunctatus, Ceutorhynchus quercicola var. erythropterus., Ceutorhynchus rudissimus, Ceutorhynchus sardeanensis, Ceutorhynchus sardeanensis diversirostris, Ceutorhynchus sartus, Ceutorhynchus sophiae, Ceutorhynchus stachydis, Ceutorhynchus striatellus, Ceutorhynchus tibialis, Ceutorhynchus timidus, Ceutorhynchus typhae, Cidnorrhinus fausti, Coeliodes ilicis, Coeliodes johannita, Coeliodes pudicus, Coeliodes rana, Coeliodes ruber, Coeliodes serratulae, Coeliodes transversealbofasciatus, Coeliodinus, Coeliodinus subrubicundus, Curculio arquata, Curculio brassicae, Curculio cardui, Curculio cruciger, Curculio grypus, Curculio invasor, Curculio maculaalba, Curculio obstrictus, Curculio punctulum, Curculio rubicundus, Curculio rugulosus, Curculio subrufus, Curculio terminatus, Datonychus arquata, Dieckmannius sexnotatus, Glocianus, Glocianus bituberculatus, Glocianus moelleri, Glocianus rudissimus, Hirtegrius, Hirtegrius setosus, Hypohypurus testaceirostris, Indozacladus cajani, Isorhynchus, Isorhynchus candidulus, Isorhynchus carinifrons, Isorhynchus porcellus, Isorhynchus squamiger, Lioxyonyx, Lioxyonyx glabirostris, Micrelus, Micrelus humilis, Micrelus polliger, Micrelus rhusii, Micrelus rinicae, Micrelus usambaricus, Microplontus fairmairii, Microplontus rugulosus, Mogulones abbreviatulus, Mogulones borraginis, Mogulones fausti, Mogulones geographicus, Mogulones javetii, Mogulones pallidicornis, Nedyus quadrimaculatus, Neoglocianus maculaalba, Neophytobius, Notegrius, Oplitoxyonyx stigma, Oprohinus libanoticus, Oreorrhynchaeus hispanicus, Parethelcus pollinarius, Paroxyonyx, Pelenomus commari, Pelenomus scoliasus, Perigasteromimus mixtus, Perigasteromimus tetracanthus, Perigasteromimus., Perioxyonyx, Phytobius leucogaster, Phytobius mixtus, Phytobius paraguayanus, Prisistus (Austroceutorhynchus), Prisistus (Ranunculiphilus), Prisistus luteitarsis, Pseudocoeliodes rubricus, Ranunculiphilus -, Ranunculiphilus (Austroceutorhynchus), Rhinoncus, Rhinoncus seniculus, Rhinoncus sparsesetosus, Rhinoncus stigma, Rhynchaenus crenatus, Rhynchaenus guttalis, Rhynchaenus nebulosus, Rhynchaenus olivaceus, Rhynchaenus paroculus, Rhynchaenus variegatulus, Rutidosoma globulus, Sirocalodes lineolaalba, Sirocalodes quercicola -, Sirocalodes sericans, Sirocalodes siculus, Stenocarus ruficornis, Thamiocolus, Thamiocolus fausti, Thamiocolus schneideri, Theodorinus hispanicus, Trichosirocalus barnevillei, Xenysmoderes helferi, Xenysmoderes infuscatus, Zacladus geranii
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Study of types and literature survey led to the discovery of 75 new synonymies and 65 new combinations. Neotypes are designated for 14 spp. as are lectotype/paralectotype designations or validations for 61 spp. Precise dates of publication of Ceutorhynchus angustus Dieckmann and Smreczynski, 1972; and that of Microplontus fairmairii (C. Brisout, 1881) are given. The authorship of: Ceutorhynchus sophiae Gyllenhal, 1837; Glocianus rotundatus (Kraatz, 1868); Rhinoncus perpendicularis form lysholmi (Schultze, 1901) are rectified. The taxonomy of Microplontus subfasciatus (Chevrolat) and of Mogulones crux (Stephens, 1831) is discussed. The current wrong spellings of Datonychus arquata (Herbst) and of Microplontus fairmairii (C. Brisout, 1881) are corrected. A selection among the alternative original spellings of Acallodes saltoides Dietz, 1896, Barioxyonyx Hustache, 1931, Ceutorhynchus erythropus (Dietz, 1896), Paroxyonyx Hustache, 1931, Pelenomus scoliasus (Dietz, 1896), Perioxyonyx Hustache, 1931, Theodorinus hispanicus Colonnelli, 1995 and Xenysmoderes infuscatus (Korotyaev, 1989) is made. Amalus alpinus Hampe, 1897 is designated as the type species of Brachiodontus Schultze, 1897; Curculio inconspectus (Herbst, 1795) is selected as the type species of Cryptoris Billberg, 1820. The following species, whose types are lost, are considered nomina dubia: Isorhynchus candidulus (Gyllenhal), Isorhynchus squamiger (Gyllenhal), Micrelus polliger (Gyllenhal) and Rhinoncus sparsesetosus Gyllenhal. The following names are revived from synonymy: Ceutorhynchus typhae (Herbst, 1795); Microplontus chrysanthemi (Gyllenhal, 1827); and Pelenomus commari (Panzer, 1794). New names are Ceutorhynchus primevus for Ceutorhynchus obliquus B. Forster; 1891 not LeConte, 1876; and Indozacladus cajani for Ceutorhynchus asperulus Faust, 1898 not Boheman, 1845.

Taxonomic name: 
Ceutorhynchinae (Alonso-Zarazaga & Lyal, 1999 (amended)), Barioxyonyx (Alonso-Zarazaga & Lyal, 1999 (amended)), Brevicoeliodes (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodinus (Alonso-Zarazaga & Lyal, 1999 (amended)), Glocianus (Alonso-Zarazaga & Lyal, 1999 (amended)), Isorhynchus (Alonso-Zarazaga & Lyal, 1999 (amended)), Micrelus (Alonso-Zarazaga & Lyal, 1999 (amended)), Paroxyonyx (Alonso-Zarazaga & Lyal, 1999 (amended)), Perioxyonyx (Alonso-Zarazaga & Lyal, 1999 (amended)), Thamiocolus (Alonso-Zarazaga & Lyal, 1999 (amended)), Aoxyonyx (Alonso-Zarazaga & Lyal, 1999 (amended)), Notegrius (Alonso-Zarazaga & Lyal, 1999 (amended)), Lioxyonyx (Alonso-Zarazaga & Lyal, 1999 (amended)), Neophytobius (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhinoncus (Alonso-Zarazaga & Lyal, 1999 (amended)), Brachiodontus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus cochleariae (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus fallax (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus granulicollis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus griseus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus sardeanensis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus tibialis (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes ruber (Alonso-Zarazaga & Lyal, 1999 (amended)), Pseudocoeliodes rubricus (Alonso-Zarazaga & Lyal, 1999 (amended)), Stenocarus ruficornis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus nanus (Alonso-Zarazaga & Lyal, 1999 (amended)), Phytobius leucogaster (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes rana (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus abchasicus var. tyli (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus insecabilis (Alonso-Zarazaga & Lyal, 1999 (amended)), Lioxyonyx glabirostris (Alonso-Zarazaga & Lyal, 1999 (amended)), Acallodes saltoides (Alonso-Zarazaga & Lyal, 1999 (amended)), Amicroplontus arcuatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceuthorhynchideus minimus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceuthorhynchidius porcellus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceuthorrhynchus cingulatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceuthorrhynchus gibbicollis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceuthorrhynchus pudicus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceuthorrhynchus sainteclairei (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus albicans (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus assimilis var. major (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus carnifrons (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus chalibaeus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus distans (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus erythropus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus gallorhenanus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus granipennis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus grenieri (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus ilvensis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus kaufmanni (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus minutus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus contractus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus moguntiacus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus nigrostriatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus obesulus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus pandellei (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus picitarsis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus pictus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus primevus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus quadripunctatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus quercicola var. erythropterus. (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus rudissimus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus sardeanensis diversirostris (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus sartus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus sophiae (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus stachydis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus striatellus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus timidus (Alonso-Zarazaga & Lyal, 1999 (amended)), Ceutorhynchus typhae (Alonso-Zarazaga & Lyal, 1999 (amended)), Cidnorrhinus fausti (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes ilicis (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes johannita (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes pudicus (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes serratulae (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodes transversealbofasciatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Coeliodinus subrubicundus (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio arquata (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio brassicae (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio cardui (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio cruciger (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio grypus (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio invasor (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio maculaalba (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio obstrictus (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio punctulum (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio rubicundus (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio rugulosus (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio subrufus (Alonso-Zarazaga & Lyal, 1999 (amended)), Curculio terminatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Datonychus arquata (Alonso-Zarazaga & Lyal, 1999 (amended)), Dieckmannius sexnotatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Glocianus bituberculatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Glocianus moelleri (Alonso-Zarazaga & Lyal, 1999 (amended)), Glocianus rudissimus (Alonso-Zarazaga & Lyal, 1999 (amended)), Hirtegrius (Alonso-Zarazaga & Lyal, 1999 (amended)), Hirtegrius setosus (Alonso-Zarazaga & Lyal, 1999 (amended)), Hypohypurus testaceirostris (Alonso-Zarazaga & Lyal, 1999 (amended)), Indozacladus cajani (Alonso-Zarazaga & Lyal, 1999 (amended)), Isorhynchus candidulus (Alonso-Zarazaga & Lyal, 1999 (amended)), Isorhynchus carinifrons (Alonso-Zarazaga & Lyal, 1999 (amended)), Isorhynchus porcellus (Alonso-Zarazaga & Lyal, 1999 (amended)), Isorhynchus squamiger (Alonso-Zarazaga & Lyal, 1999 (amended)), Micrelus humilis (Alonso-Zarazaga & Lyal, 1999 (amended)), Micrelus polliger (Alonso-Zarazaga & Lyal, 1999 (amended)), Micrelus rhusii (Alonso-Zarazaga & Lyal, 1999 (amended)), Micrelus rinicae (Alonso-Zarazaga & Lyal, 1999 (amended)), Micrelus usambaricus (Alonso-Zarazaga & Lyal, 1999 (amended)), Microplontus fairmairii (Alonso-Zarazaga & Lyal, 1999 (amended)), Microplontus rugulosus (Alonso-Zarazaga & Lyal, 1999 (amended)), Mogulones abbreviatulus (Alonso-Zarazaga & Lyal, 1999 (amended)), Mogulones borraginis (Alonso-Zarazaga & Lyal, 1999 (amended)), Mogulones fausti (Alonso-Zarazaga & Lyal, 1999 (amended)), Mogulones geographicus (Alonso-Zarazaga & Lyal, 1999 (amended)), Mogulones javetii (Alonso-Zarazaga & Lyal, 1999 (amended)), Mogulones pallidicornis (Alonso-Zarazaga & Lyal, 1999 (amended)), Nedyus quadrimaculatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Neoglocianus maculaalba (Alonso-Zarazaga & Lyal, 1999 (amended)), Oplitoxyonyx stigma (Alonso-Zarazaga & Lyal, 1999 (amended)), Oprohinus libanoticus (Alonso-Zarazaga & Lyal, 1999 (amended)), Oreorrhynchaeus hispanicus (Alonso-Zarazaga & Lyal, 1999 (amended)), Parethelcus pollinarius (Alonso-Zarazaga & Lyal, 1999 (amended)), Pelenomus commari (Alonso-Zarazaga & Lyal, 1999 (amended)), Pelenomus scoliasus (Alonso-Zarazaga & Lyal, 1999 (amended)), Perigasteromimus. (Alonso-Zarazaga & Lyal, 1999 (amended)), Perigasteromimus mixtus (Alonso-Zarazaga & Lyal, 1999 (amended)), Perigasteromimus tetracanthus (Alonso-Zarazaga & Lyal, 1999 (amended)), Phytobius mixtus (Alonso-Zarazaga & Lyal, 1999 (amended)), Phytobius paraguayanus (Alonso-Zarazaga & Lyal, 1999 (amended)), Prisistus (Austroceutorhynchus) (Alonso-Zarazaga & Lyal, 1999 (amended)), Prisistus (Ranunculiphilus) (Alonso-Zarazaga & Lyal, 1999 (amended)), Prisistus luteitarsis (Alonso-Zarazaga & Lyal, 1999 (amended)), Ranunculiphilus - (Alonso-Zarazaga & Lyal, 1999 (amended)), Ranunculiphilus (Austroceutorhynchus) (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhinoncus seniculus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhinoncus sparsesetosus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhinoncus stigma (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhynchaenus crenatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhynchaenus guttalis (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhynchaenus nebulosus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhynchaenus olivaceus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhynchaenus paroculus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rhynchaenus variegatulus (Alonso-Zarazaga & Lyal, 1999 (amended)), Rutidosoma globulus (Alonso-Zarazaga & Lyal, 1999 (amended)), Sirocalodes lineolaalba (Alonso-Zarazaga & Lyal, 1999 (amended)), Sirocalodes quercicola - (Alonso-Zarazaga & Lyal, 1999 (amended)), Sirocalodes sericans (Alonso-Zarazaga & Lyal, 1999 (amended)), Sirocalodes siculus (Alonso-Zarazaga & Lyal, 1999 (amended)), Thamiocolus fausti (Alonso-Zarazaga & Lyal, 1999 (amended)), Thamiocolus schneideri (Alonso-Zarazaga & Lyal, 1999 (amended)), Theodorinus hispanicus (Alonso-Zarazaga & Lyal, 1999 (amended)), Trichosirocalus barnevillei (Alonso-Zarazaga & Lyal, 1999 (amended)), Xenysmoderes helferi (Alonso-Zarazaga & Lyal, 1999 (amended)), Xenysmoderes infuscatus (Alonso-Zarazaga & Lyal, 1999 (amended)), Zacladus geranii (Alonso-Zarazaga & Lyal, 1999 (amended))
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith