The classification of weevils (Coleoptera: Curculionoidea) has been in flux for decades. The problem is partly due to the difficulties in recognizing monophyletic units among the higher taxa, and partly because of their poor characterization. With the gradual clarification of relationships between families and subfamilies a difference in viewpoints has emerged between those that favour fewer larger taxa and those who recognize a more atomized classification. The two most recent views are given as separate trees on the left-hand bar of this website. The classification at the top is amended from that of Alonso-Zarazaga & Lyal, 1999, which broadly follows that of Thompson, 1992. It includes all genera, since they placed them in their catalogue, and it used as the default classification on this site because of the detail possible. The alternative is that presented by Oberprieler, Marvaldi & Anderson, 2007, building both on their earlier work and that of Kuschel, 1995. Oberprieler et al’s classification is not complete, but comments can be made on pages linked to any of the taxa shown.
There have been a number of excellent studies and critiques of weevil higher classification and diversity in the past few years. Most if not all papers can be found in the bibliography of this website, but important works include:
Anderson, R.S. 1995. An evolutionary perspective on diversity
in Curculionoidea. Mem. Ent Soc Wash 14:103-114.
Cognato, A.I. & Grimaldi, D. (2009) 100 million years of
morphological conservation in bark beetles (Coleoptera: Curculionidae: Scolytinae).
Systematic Entomology, 34, 93–100.
Farrell, B.D. 1998. "Inordinate fondness" explained:
Why are there so many beetles? Science
281:553-557. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Farrell_1998.pdf
Farrell, B.D.; Sequeira, A.S.; O'Meara, B.; Normark, B.B.; Chung, J.H.; Jordal, B.H. 2001.
The evolution of agriculture in beetles (Curculionidae: Scolytinae and
Platypodinae). Evolution
55:2011-2027. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Farrell_etal_2001.pdf
Farrell, B.D.; Sequeira, A.S. 2004. Evolutionary rates in
the adaptive radiation of beetles on plants. Evolution 58:1984-2001. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Farrell&Seq2004.pdf
Franz, N.M. & Engel, M.S., 2010. Can higher-level phylogenies of weevils
explain their evolutionary success? A critical review. Systematic
Entomology, 35, 597–606. DOI:
10.1111/j.1365-3113.2010.00534.x
Hunsdoerfer, A.K.; Rheinheimer, J.; Wink, M. 2009.
Towards the phylogeny of the Curculionoidea (Coleoptera):
Reconstructions from mitochondrial and nuclear ribosomal DNA sequences. Zoologischer
Anzeiger 248 (2009) 9-31
Hunt, T.; Bergsten, B.; Levkanicova, Z.; Papadopoulou, A.; St
John, O.; Wild, R.; Hammond, P.M.; Ahrens, D.; Balke, M.; Caterhino, M.S.;
Gomez-Zurita, J.; Ribera, I.; Barraclough, T.G.; Bocakova, M.; Bocak, L.;
Vogler, A.P. 2007. A comprehensive phylogeny of beetles reveals the
evolutionary origins of a superradiation. Science
318:1913-1916.
Jordal, B.H.; Normark, B.B.; Farrell, B.D. 2000. Evolutionary
radiation of an inbreeding haplodiploid beetle lineage (Curculionidae, Scolytinae).
Biological Journal of the Linnean Society
71:483-499. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/jordaletal2000.pdf
Jordal, B.H., 2007. Reconstructing
the Phylogeny of Scolytinae and Close Allies: Major Obstacles and Prospects for
a Solution. USDA Forest Service
Proceedings RMRS-P-45. 1-9 http://www.fs.fed.us/rm/pubs/rmrs_p045/rmrs_p045_003_009.pdf
Jordal, B.J., Sequeira, A.S. & Cognato, A.I., 2011. The age and phylogeny of wood boring weevils
and the origin of subsociality. Molecular Phylogenetics and Evolution. 59(3):708-24.
Epub 2011 Mar 22.
Kuschel, G. 1995. A phylogenetic classification of Curculionoidea
to families and subfamilies. Mem Ent Soc
Wash 14:5-33.
Kuschel, G.; Leschen, R.A.B.; Zimmerman, E.C. 2000.
Platypodidae under scrutiny. Invertebrate
Taxonomy 14:771-806.
Lefevre, C.; Charles, H.; Vallier, A.; Delobel, B.; Farrell,
B.D.; Heddi, A. 2004. Endosymbiont phlyogenesis in the Dryophthoridae weevils:
Evidence for bacterial replacement. Molecular
Biology and Evolution 21: 965-973. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Lefevre_et_al_2004.pdf
Marvaldi, A.E. 1997. Higher level phylogeny of Curculionidae
(Coleoptera: Curculionoidea) based mainly on larval characters, with special reference
to broad-nosed weevils. Cladistics
13:285-312. http://www.cricyt.edu.ar/entomologia/Articulos/Cladistics13(1997).pdf
Marvaldi, A.E., 2003, Key to larvae of the South American subfamilies
of weevils (Coleoptera, Curculionoidea).
Revista Chilena de Historia
Natural, 76: 603-612.
Marvaldi, A.E. 2005. Larval morphology and biology of oxycorynine
weevils and the higher phylogeny of the Belidae (Coleoptera, Curculionoidea). Zoologica Scripta 34:37-48.
Marvaldi, A.E.; Duckett, C.N.; Kjer, K.M.; Gillespie, J.J. 2009.
Structural alignment of 18S and 28S rDNA sequences provides insights in to
phylogeny of Phytophaga (Coleoptera: Curculionoidea and Chrysomeloidea). Zoologica Scripta 38:63-77.
Marvaldi, A.E.; Lanteri, A.A., 2005. Key to higher taxa of South American weevils
based on adult characters (Coleoptera, Curculionoidea). Revista
Chilena de Historia Natural. 78:
65-87. http://www.scielo.cl/pdf/rchnat/v78n1/art06.pdf
Marvaldi, A.E.; Morrone, J.J. 2000. Phylogenetic systematics
of weevils (Coleoptera: Curculionoidea): A reappraisal based on larval and
adult morphology. Insect Syst Evol
31:43-58.
Marvaldi, A.E.; Oberprieler, R.G.; Lyal, C.H.C.; Bradbury, T.;
Anderson, R.S. 2006. Phylogeny of the Oxycoryninae s.l. (Coleoptera Phytophaga)
and evolution of plant-weevil interactions. Invertebr
Syst 20:447-476.
Marvaldi, A.E.; Sequeira, A.S.; O'Brien, C.W.; Farrell, B.D.
2002. Molecular and morphological phylogenetics of weevils (Coleoptera:
Curculionoidea): Do niche shifts accompany diversification? Systematic Biology 51:761-785. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Marvaldietal2002.pdf
McKenna, D.D.; Farrell, B.D. 2006, Tropical forests are both
evolutionary cradles and museums of leaf beetle diversity. PNAS,
103 (29): 10947-10951 (July 18, 2006) http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/McKenna_Farrell_2006.pdf
McKenna, D.D.; Sequeira, A.S.; Marvaldi, A.E.; Farrell, B.D.,
2009, Temporal lags and overlap in the
diversification of weevils and flowering plants. PNAS
Early Edition, www.pnas.org_cgi_doi_10.1073_pnas.0810618106(1-6)
Morrone, J.J., 2000. Mexican weevils (Coleoptera: Curculionoidea):
a preliminary key to families and subfamilies.
Acta Zoo. Mex. (n.s.)
80:131-141
Normark, B.B.; Jordal, B.H.; Farrell, B.D. 1999. Origin of a
haplodiploid beetle lineage. Proceedings
of the Royal Society Biological Series B 266:2253-2259. http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Normark_etal_1999.pdf
Oberprieler, R.G.; Marvaldi, A.E.; Anderson, R.S. 2007. Weevils,
weevils, weevils everywhere. Zootaxa
1668:491-520.
Sequeira, A.S.; Farrell, B.D. 2001. Evolutionary origins of
Gondwanan interactions: How old are Araucaria beetle herbivores? Biological Journal of the Linnean Society
74:459-474 http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/Sequeira_Farrell_2001.pdf
Sequeira, A.S.; Normark, B.B.; Farrell, B.D. 2000. Evolutionary
assembly of the conifer fauna: Distinguishing ancient from recent associations
in bark beetles. Proceedings of the Royal
Society Biological Series B 267:2359-2366.
http://insects.oeb.harvard.edu/farrell_lab/people/farrell/papers/sequeira_2000.pdf
Thompson, R.T. 1992. Observations on the morphology and classification
of weevils (Coleoptera: Curculionoidea) with a key to the major groups. J Nat Hist 26:835-891.